Senior Lecturer & Manager - Central Science Consultancy Bureau , College Of Natural And Applied Sciences

Ph.D (Zoology); Specialization in Ornithology, University of Dar es Salaam, Tanzania.


Dr. Nyamkinda has been teaching the following courses:

  • Chordate Zoology
  • Ornithology
  • Animal behaviour
  • Reproduction in wild mammals
  • Biodiversity Conservation


Research interests:  He is very passionate in researching birds and bats of Tanzania, and elsewhere in Africa.


  1. Promoting parents and teachers awareness on the psychological wellbeing of primary school children in boarding schools in Tanzania.
  2. Molecular epidemiology and antimicrobial resistance of Campylobacter species causing diarrhoea in Dar es Salaam.
  3. Wildlife Management Areas (WMAS) and Community Wellbeing in Tanzania.


  1. Werema, C. (Accepted). Short-term responses of birds to prescribed fire in Combretum woodland in Tarangire National Park, Northern Tanzania. African Journal of Ecology (Accepted for publication in African Journal of Ecology).
  2. Werema, C. (2020).Seasonal abundance and distribution of Red-capped Robin Chat Cossypha natalensis in the Eastern Arc Mountains, Tanzania. Ostrich 91(2): 129-133, DOI: 10.2989/00306525.2020.1752838
  3. Werema, C., Senzota, R.B.M. and Mohamed, N. (2020). Avifaunal Biodiversity in Peril: Pre- and Post-Vegetation Clearance Survey of Avifauna in a Fragmented Thicket in an Urban Landscape, Dar Es Salaam City, Tanzania. Journal of East African Natural History 108(2): 65-80.
  4. Summay, G., Lyakurwa, K., Lyakurwa J., Werema, C., Tito, L. Raymond, O. (2020). The Reptiles of Mkomazi National Park, Tanzania an Updated Checklist with Some Aspects of Biogeography. Journal of East African Natural History 108(2):81-98.
  5. Werema, C. and Msuya, C.A. (2019).Understorey Bird Communities 8 and 18 Years after River Diversion in Kihansi Gorge, Udzungwa Mountains in the Eastern Arc Mountains, Tanzania. Tanzania Journal of Science 45(4): 661-671.
  6. Werema, C. (2017). Preliminary comparison of birds inhabiting exotic Acacia and native bushland habitats in semi-arid east-central Tanzania. Scopus 37(2): 32-37.
  7. Werema, C., Nahonyo, C.L. and Kibaja, M. (2017). The role of kopjes in bird species’ conservation within an agricultural matrix west of the Greater Serengeti Ecosystem, Tanzania. Scopus 37(2): 8-23.
  8. Shirima, J.D. and Werema, C. (2017). Effect of habitat fragmentation on diversity and abundance of nesting birds in an urban landscape: the case of Mwalimu Nyerere campus thickets, University of Dar es Salaam, Tanzania. Tanzania Journal of Science 43(1): 14-22.
  9. Werema, C., Howell, K.M. and Ndangalasi, H.J. (2016). Use of remnant forest fragments by seasonal altitudinal migrating forest birds in the Uluguru Mountains, Tanzania: A conservation priority. Ostrich 87(3) 255-262.
  10. Werema, C. (2016). Seasonal variation in understorey bird species diversity and abundance in the Uluguru Nature Reserve, Tanzania. African Journal of Ecology. 54(3): 299-307. doi: 10.1111/aje.12219.
  11. Werema, C., Howell, K.M. and Ndangalasi, H.J. (2016). Kimboza Forest Reserve, Tanzania: an important cold season refugium for altitudinal migrating birds. African Journal of Ecology.54(3): 375-378. DOI: 10.1111/aje.12327.
  12. Werema, C. (2016). Tanzania Red-billed Hornbill Tockus ruahae feeding on a Giant Water Bug. Scopus 36(2): 52-53.
  13. Werema, C., Howell, K.M., Msuya, C.A., Sinclair, J. and Macha, A. (2016) Birds of Golden Pride Project area, Nzega District, central Tanzania: an evaluation of recolonisation of rehabilitated areas. Scopus 36(2): 26-36.
  14. Werema, C. (2016). Seasonal elevational movements of Eastern Olive Sunbird Cyanomitra olivacea (A. Smith) in the Uluguru Mountains, Tanzania. Ostrich 87(2): 189-192.
  15. McEntee, J.P., Peñalba, J.V., Werema, C., Mulungu, E., Mbilinyi, M., Hansen, L., Fjeldsa, J. and Bowie, R.C.K. (2016). Social selection parapatry in an Afrotropical sunbird. Evolution. doi:10.1111/evo.12950.
  16. Werema, C. and Howell, K.M. (2016). Seasonal variation in diversity and abundance of understorey birds in Bunduki Forest Reserve, Tanzania: evaluating the conservation value of a plantation forest. Ostrich 87(1): 89-93.
  17. Werema, C. and Nahonyo, C.L. (2015). Preliminary observations of the birds of Songo Songo Island, Tanzania. Scopus 35: 39-43.
  18. Werema, C. (2014). Effects of fire on understorey birds in Kimboza Forest Reserve in the eastern foothills of the Uluguru Mountains, Tanzania. African Journal of Ecology. doi: 10.1111/aje.12183.
  19. Werema, C. (2015). Seasonal elevational movements of the Little Greenbul Andropadus virens in the Uluguru Mountains, Tanzania. African Journal of Ecology 53: 253–256. doi: 10.1111/aje.12190.
  20. Werema, C., McEntee, J.P., Mulungu, E. and Mbilinyi, M. (2013). Preliminary observations on the birds of Ikokoto forest, Udzungwa Mountains, Tanzania. Scopus 32: 19-26.

Books and Book chapter

  1. John, J, Kohi, E., Park, J., Nkwabi, A, Choi, C., Werema, C., Hur, W., Kim, H. and Lee, J. (2019). Birds of Western Tanzania (Gombe, Mahale, Rubondo, Saanane, Malagarasi, Ntakata and Minziro. National Institute of Biological Resource, South Korea.
  2. Gereau, R.E., Kariuki, M., Ndang’ng’a P. K, Werema, C. and Muoria, P. (2014). Biodiversity Status and Trends in the Eastern Arc Mountains and Coastal Forests of Kenya and Tanzania Region, 2008-2013. Nairobi: BirdLife Internatinal-Africa Partnership Secretariat.
  3. Mlingwa, C., Ntongani, W. and Werema, C. (2012). Checklist of Tanzanian Birds. Pp. 65-113 in Gideon, H., Nyinondi, P, and Oyema, G. Checklist of Tanzanian Species. InterPress of Tanzania, Dar es Salaam.